ScottB:H2Andy's link says pretty much the same thing, but here are my two references (no links, sorry)
ANDI International Recreational Diving with Closed Circuit Rebreathers: ISC Megalodon Rebreather is where I got my information.
Chapter 2 Page 13:
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Thanks for that. Andy's reference was a little vague on the cause but vasodilation makes sense.
Funnily enough I had already ordered the manual you quoted and it should be here "any day now"