I'm confused. We see nudibranch mating all the time over here and I guaranty that they do it a heck of a lot more than once a year. We also see lots of egg masses and the nudis laying them and it never happens like you describe. I'm not saying you are wrong but this sequence may be fairly unique to this species. Have you been able to identify it?
The species is a Florida Regal Sea Goddess... the only species of nudi we see here. And while very common, we don't see egg masses, but that might be because they hide them.
This is a very seasonal environment here.. going from water temperatures less than 50 F to over 85 F (less than 10 C to over 30 C)..life tends to revolve around temperature.
Their food (they are algae eaters) only grows during some of the year. Short days and low vis are the norm during the winter months, and these guys typically live around 30 meters. They only clump together in the May time frame...Water has warmed, days are much longer and the water tends to be clearer.
Those images are time stamped, and the first 4 represent around 2 minutes of time (feel free to check yourself, I left all the data on the images. )
As you see them mate all the time, would you be nice enough to post images of what they do do? As I have no idea, if it is different, how different it is.
There should be no surprise in their mating being seasonal, Sea Turtles do the same thing... those in say the great barrier reef mate and lay eggs all the time.... ones living in seasonal areas only do it once a year....Our trees do that also.
I have no idea how they know when to clump.. how they manage to pick mates, or even how they managed to know what to do. Seems to me a pretty complex process, and my take of sea slugs is that they are not the brightest animal in the ocean. But if you look at the timing of those images, the passing the lump back and forth took place fairly quickly. I am not surprised at all that warm water ones do things differently.