Hank49:
If I'm correct in thinking I understand this, literally at an instant when a mother gave birth, (with different species throughout time) she gave birth to a different species. Because at some point, the DNA compatibility went from, "can breed" to "no can breed" (to original parent stocks which hadn't mutated or changed). There would be no "maybe". Correct? This is what I see being questioned. At some point this had to have happened.
put it this way:
a bunch of wolves break off the rest of the wolves and start breeding amongst themslves. they happen to have a larger percentage of large wolves, so they start getting bigger. then it gets really cold, and those with thick fur survive more and so reproduce more.
so after a while, you have larger, furrier wolves.
at this point, both "tribes" can mate and have children, which look like a cross between the two parents (where dogs are today).
due to the extra cold, they have to change their hunting tactics and those who can hunt by themselves rather than in a pack survive and breed.
thus, you end up with larger, furrier, solitary hunting wolves. since the new "animal" is solitary now, it has real trouble relating to the pack-mentality of the other wolves, so they really don't have much contact anymore. inter-breeding pretty much ends.
still, it is cold, and there's not much to hunt. some of the "new" wolves start swimming in streams after prey. those with bigger paws do better, so their paws get larger as this gene is passed down to other generations.
then, the fur of some of the wolves being oilier allows them to survive better. you end up with very oily-furred wolves that can thrive in the new environment.
it has been such a gradual change that at any time, a parent and an off-spring can mate, but if you look at what you started with:
wolves that hunt in packs
and what you ended up with:
large, solitary, slick-hared animals, with huge paws, that swim and hunt by themselves
you have two rather different species already, their DNA different enough that they probably wouldn't be able to have succesful offpsring
if you keep adding small changes over time, you can end up with any of a number of species